Bachelor groups in the summer, in areas where cows don't frequent, is common behavior for elk.
As the rut approaches, things will change as the bulls become less tolerant of each others company. In the first part of September they will often stay in the same location as their summer hangout, but spread out a little to avoid each other. Wallows can be especially important at this time, where bulls can cool off and relieve some of the rutting tension that is starting to build. If you are hunting with a bow early in September, staying relatively close to the summer range is your best bet to find some of those bachelors. Calling can be effective, but be patient. Early in September bulls may not be that vocal but will often check out a caller by approaching silently to see who is making all the noise.
In the first part of September, some of the younger bulls may drift towards and start accompanying the cow herds. This may be close to the bachelor's summer range or a fair distance away. Often cows summer in areas with the best feed, and the bulls are in more marginal habitat, so the distance apart can depend on the local area. Towards the middle of September, and into early October, the mature bulls will move towards the cow herds and take over from the younger bulls.
Other variables can influence these movements and ranges, including feed quality, predators, human pressure, cows....so nothing is for certain. My experience with this is primarily in mountain habitat, and the 400 WMU series you mention fits that description. Hope that helps and let us know how you make out. Best of luck.
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